During the period of the primary contact time,there are series of recognition between the Fungal pathogens and the plant cell,which include physical and biochemical recognition.The results affect the infection directly.In the process of penetrating plant cell walls,there is not only an emzymolysis to plant cell wall,but also a mechanical squeeze to the cell wall.The latter almost trigger cell-associated defense response,such as extracellular hydrogen peroxide generation and callose doposition The complete cell wall-related defence response will be induced by mechanical signal and chemical signal together.Being to with the mammalian cell,in plant cell wall intergrin-like proteins have response to perceive the mechanical signal,which is via RGD(Arg-Gly-Asp)motif,and the plasma membrane-cell wall adhesion is required for expression of plant defense responses during fungal penetration.